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This repository contains all R-scripts and respective model output data needed to reproduce the methods, results and figures of the manuscript Browsing herbivores improve the state and functioning of savannas: a model assessment of alternative land use strategies by Katja Irob, Niels Blaum, Selina Baldauf, Leon Kerger, Ben Strohbach, Angelina Kanduvarisa, Dirk Lohmann and Britta Tietjen. For any questions regarding content in this repository please contact Katja Irob.

R-Scripts

R-Scripts to reproduce all figures and statistical procedures are provided in the folder Analysis and the corresponding data in the Data folder. Scripts for the evaluation of the sensitivity analysis output and plant species parameterisation are in the folder Sensitivity, corresponding sensitivity output data in the folder Input. If you want to reproduce the results please download the repository and run the code with R (most conveniently in R-Studio). For any questions regarding the R-Scripts please contact Katja Irob.

Script Description
Analysis/Cover.R Script to visualise and analyse the meta-PFT cover time series for the land use scenarios used in the main part of the manuscript
Analysis/Cover_appendix.R Script to create the meta-PFT cover time series for all other land use scenarios presented in the appendix
Analysis/Composition.R Script to visualise and analyse the strategy-type composition of the last 20 years of simulation for each scenario used in the main part of the manuscript
Analysis/Composition_appendix.R Script to visualise the strategy-type composition of the last 20 years of simulation for additional scenarios presented in the appendix
Analysis/Richness_Evenness.R Generation of richness and evenness values, visualisation and analysation of the data of the last 20 years of simulation for each scenario used in the main part of the manuscript
Analysis/Richness_Evenness_appendix.R Generation of richness and evenness values and visualisation of the data of the last 20 years of simulation for additional scenarios presented in the appendix
Analysis/EFA_Clusters.R Exploratory factor analysis to determine clusters depending on plant strategies. Additional calculation of functional dispersion (FDis) and visualisation of PFT clusters including individual abundance and distance to centroid for the main scenarios
Analysis/Water.R Calculation of T/ET, visualisation of T/ET and soil moisture and statistical evaluation for the main scenarios
Analysis/Water_appendix.R Calculation of T/ET, visualisation of T/ET and soil moisture for all additional scenarios
Sensitivity/Parameterisation_sensitivity_browse.R Merging sensitivity output for all parameters and climate repetitions, determining parameter value at desired cover change based on linear regression analysis, visualisation of output for browsing scenarios
Sensitivity/Parameterisation_sensitivity_graze.R Merging sensitivity output for all parameters and climate repetitions, determining parameter value at desired cover change based on linear regression analysis, visualisation of output for grazing scenarios

Folder /Data

Folder /Results + /Appendix

This folder contains model output files for grazing and browsing scenarios for the stocking rates 20 ha/LSU and 40 ha/LSU. The filename e.g. "EH_100years_yearly_EH_SR20browse_climrep-1.txt" refers to the site "Etosha Heights" (EH), 100 years simulation time and yearly output, the stocking rate (SR) 20 ha/LSU of browsers and the climate repetition 1 (of 30). /Appendix additionally contains the result files for the stocking rates 10, 20, 30 and 50. The files contain the complete model output and are condensed to the variables of interest using the R scripts in /Analysis. Only the output variables used in this study are explained below.

File Description
EH_100years_yearly_EH_SR20/40browse/graze_climrep-x.txt EcoHyD simulation output for 100 years based on environmental conditions at Etosha Heights, Namibia.
Year: year from 0-99
meanGcover_x: mean perennial grass cover of strategy types 0-9 [-]
meanScover_x: mean shrub cover of strategy types 0-11 [-]
meanAcover_x: mean annual grass cover of base-type [-]
meanGtotalcover: total perennial grass cover [-]
meanStotalcover: total shrub cover [-]
meanAtotalcover: total annual grass cover [-]
Annualevaporation: total annual evaporation [Vol%]
AnnualtranspirationL1: total annual plant transpiration of layer 1 [Vol%]
ML1: soil moisture in layer 1 during rainy season [Vol%]

Folder /Matrices_for_EFA

Matrices used for calculation of functional dispersion as part of the factor analysis in script /Analysis/EFA_Clusters.R.

File Description
cattle_matrix.txt Matrix containing abundance data for all plant strategy-types (columns) for each grazing scenario (rows).
trait_matrix_cattle.txt Matrix containing plant strategy-type trait values used for parameterisation in grazing scenarios.
wildlife_matrix.txt Matrix containing abundance data for all plant strategy-types (columns) for each browsing scenario (rows).
trait_matrix_wildlife.txt Matrix containing plant strategy-type trait values used for parameterisation in browsing scenarios.

Folder /Model

Model source files can be found in /Source. To reproduce examples, download repository and keep folder structure. Output files are stored in /Results.

File Description
compile.sh Example bash script to compile and run the model.

Folder /Parameters

File Description
elevation_30_EH.txt Digital elevation map in 30 by 30 grid based on study site at Etosha Heights. [m]
modelparameters_control.txt General configuration parameters for model EcoHyD.
Site: name of site [-]
Latitude: latitude of the site [°S]
simYears: total simulation time [years]
xsize: x length of the grid (number of x-cells) [-]
ysize: y length of the grid (number of y-cells) [-]
cellsize: size of a single cell [m]
vegTimeStep: number of days after which vegetation is updated [days]
modelscenarios.txt Scenarios, climate repetitions and repetitions are determined here.
Input files will be chosen and output files will be coded based on IDs and repetitions you indicate here.
soilparameters_EH.txt Soil parameters for loamy sand, calibrated for Etosha Heights.
vegetationparameters_ScenarioName.txt General vegetation parameters and specific parameters for the PFTs shrubs, perennial and annual grasses. Sub-types are generated here.

Folder /Weather

Climate input data for EcoHyD. Precipitation and temperature time series generated in NamRain based on TRMM series.

File Description
EH_100years_control_climrep-1-30.txt Climate files at hourly resolution in Y (year), M (month), D (day), H (hour) format.
Rain: Hourly precipitation amount [mm]
Temp: Hourly temperature [°C]

Folder R

File Description
Utility.R R script to extract often repeating functions from, e.g. reading in data, renaming species, etc.

Folder /Sensitivity

Folder /Input

The folders Browse and Graze hold the single result files of the sensitivity analysis for grazing and browsing respectively for both perennial grasses and shrubs. The parameterisation is based on this output.

File Description
/Browse/perennials_sensoutput_ParameterName.txt Cover change of perennial grasses in response to parameter value changes in browsing scenario for the parameters: biomass production (GR), mortality (MR), palatability (GP), defense (GL), competitive strength for water (UR), water needed for establishment (ES) and resistance to drought (WP).
/Browse/shrubss_sensoutput_ParameterName.txt Cover change of shrubs in response to parameter value changes in browsing scenario for the parameters: biomass production (GR), mortality (MR), palatability (GP), defense (GL), competitive strength for water (UR), water needed for establishment (ES) and resistance to drought (WP).
/Graze/perennials_sensoutput_ParameterName.txt Cover change of perennial grasses in response to parameter value changes in grazing scenario for the parameters: biomass production (GR), mortality (MR), palatability (GP), defense (GL), competitive strength for water (UR), water needed for establishment (ES) and resistance to drought (WP).
/Graze/shrubss_sensoutput_ParameterName.txt Cover change of shrubs in response to parameter value changes in grazing scenario for the parameters: biomass production (GR), mortality (MR), palatability (GP), defense (GL), competitive strength for water (UR), water needed for establishment (ES) and resistance to drought (WP).

Folder /Parms

File Description
Sens_output_parameter_browsing_na.txt Standard parameter values of the base-types of perennial grasses and shrubs used as reference for parameterisation of strategy types in browsing scenarios. Included are the parameters biomass production (GR), mortality (MR), palatability (GP), defense (GL), competitive strength for water (UR), water needed for establishment (ES) and resistance to drought (WP).
Sens_output_parameter_grazing_nc.txt Standard parameter values of the base-types of perennial grasses and shrubs used as reference for parameterisation of strategy types in grazing scenarios. Included are the parameters biomass production (GR), mortality (MR), palatability (GP), defense (GL), competitive strength for water (UR), water needed for establishment (ES) and resistance to drought (WP).

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